Taxonomic notes on Euptychia modestaButler, 1867,Neonympha alcinoe C. Felder & R. Felder, 1867 and Euptychiapamela Hayward, 1957(Lepidoptera: Nymphalidae: Satyrinae), with descriptionsof three new genera, five new species and two new subspecies
Notas taxonómicas sobre Euptychia modesta Butler, 1867, Neonympha alcinoe C. Felder & R. Felder, 1867 y Euptychia pamela Hayward, 1957(Lepidoptera: Nymphalidae: Satyrinae), con descripciones de tres géneros nuevos, cinco especies y dos subespecies nuevas de Centroamérica y Suramérica
Abstract
The identity, status and taxonomic position of seven species of Neotropical satyrine butterflies are discussed and defined: Euptychia modesta Butler, 1867, Euptychia nebulosa Butler, 1867, Neonympha alcinoe C. Felder & R. Felder, 1867, Eup-tychia mimas Godman, 1905, Euptychia mimula Hayward, 1954, Euptychia pamela Hayward, 1957 and Cissia gomeziSinger, DeVries & Ehrlich, 1983. These taxa lately remained in genera such as Cissia Doubleday, 1848, MagneuptychiaForster, 1964 and Yphthimoides Forster, 1964. Based on the particular combination of selected morphological characters and comparative DNA analysis of these and other evidently related species, three new genera (with seven new combina-tions), five new species and two new subspecies are proposed, diagnosed and described: Modestia Viloria & Benmesbah, gen. nov., with four taxa, M. modesta (Butler, 1867), comb. nov., M. gomezi (Singer, DeVries & Ehrlich, 1983), comb. nov., M. remypignoux Benmesbah & Viloria, sp. nov. and M. remypignoux shueyi Benmesbah & Viloria, ssp. nov.; Ma-laveria Viloria & Benmesbah, gen. nov., with nine taxa, M. alcinoe (C. Felder & R. Felder, 1867), comb. nov., M. nebu-losa (Butler, 1867), comb. nov., M. mimas (Godman, 1905), comb. nov., M. ballofi Benmesbah & Viloria, sp. nov., M. rodriguezi Benmesbah & Viloria, sp. nov., M. rodriguezi risaralda Benmesbah & Viloria, ssp. nov., M. bottoi Benmesbah & Viloria, sp. nov., M. duponti Benmesbah & Murienne, sp. nov. and M. mimula (Hayward, 1954), comb. nov.; and Koutalina Viloria & Murienne, gen. nov., for a single known species, K. pamela (Hayward, 1957), comb. nov. Lectotypes of Euptychia modesta,Neonympha alcinoe and Neonympha sylvina C. Felder & R. Felder, 1867, are designated. Variability in morpholog y and aspect of the male genitalia is discussed, as well as number of pupils in ocelli and the occasional pres-ence of supernumerary anal ocellus on ventral hindwing.
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